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In mosquitoes and fruit flies, sound causes fine antennal hairs to quiver. Those detectors occur throughout the insect body but evolution typically only modified a single pair - apparently, almost any pair - to perceive the airborne vibrations generated by sound.įrom there on, each new attempt to forge ears went even further in its own direction as other structures were co-opted and reconfigured to capture, amplify and filter sound, extract the relevant information and convey it to the nervous system. This anywhere-goes approach might seem a little weird but there’s a simple explanation: In every case where an insect ear evolved, the starting point was an existing sensory organ: a stretch detector that monitors tiny vibrations when neighboring body segments move. Hearing has evolved at least 20 times in insects, leading to ears in an astonishing number of different locations, as shown on this image of a generalized insect. Praying mantises have a single, “cyclopean” ear in the middle of their chest. The bladder grasshopper has an abundance of ears with six pairs along the sides of its abdomen. Among moths and butterflies, ears crop up practically anywhere, even on mouthparts. Location is the most obvious difference between one insect’s ears and another’s: There are ears on antennae (mosquitoes and fruit flies), forelegs (crickets and katydids), wings (lacewings), abdomen (cicadas, grasshoppers and locusts) and on what passes for a “neck” (parasitic flies). All told, insect ears arose more than 20 separate times, a sure-fire recipe for variety. The 350,000 species of that most dazzlingly diverse group, the beetles, are almost all deaf, yet the few that have ears acquired them through two separate lines of evolution. Of the 30 major insect orders, nine (at last count) include some that hear, and hearing has evolved more than once in some orders - at least six times among butterflies and moths. These ancestral insects went on to diversify into more than 900,000 species, and while most remain as deaf as their ancestors, some gained the means to hear. When insects first appeared some 400 million years ago, they were deaf, Göpfert tells me. And thanks to some of this newfound knowledge, and an assortment of fossil insects, there’s even the tantalizing prospect of being able to eavesdrop on the ancient past, adding a new dimension to our understanding of the life and times of some long-vanished animals. Sensory biologists, acoustics experts and geneticists are working together to pin down how they all work, how and when they evolved, and why. But with the advent of new tools and technologies, ever more examples are coming to light. Most are hard to spot, if not invisible, and in many cases insects produce and sense sounds so far beyond our own range that we overlooked their abilities entirely. Amazing though katydid ears are, he tells me, they’re just one of many with astonishing capabilities: Evolution has made so many attempts at shaping ears, the result is a huge diversity of structures and mechanisms. Most moths have ears tuned to the frequencies used by bats.ĬREDIT: AVALON / PHOTOSHOT LICENSE / ALAMY STOCK PHOTOĬuriosity piqued, I call neurobiologist Martin Göpfert at the University of Göttingen in Germany, who studies hearing in the fruit fly Drosophila melanogaster. Yet obviously, some must hear: The summer air is filled with the trills, chirps and clicks of lovelorn crickets and grasshoppers, cicadas and katydids, all trying to attract a mate.Ī greater horseshoe bat hunts a moth. The appearance of bats that hunt with the aid of ultrasonic sonar drove the evolution of hearing in many moths and other night-flying insects. Insect eyes and antennae stand out, but ears? Even the eagle-eyed could be forgiven for wondering if insects have them. I’d never given much thought to insect ears until now. One Australian katydid has capitalized on its auditory prowess to capture prey in a very devious way: It lures male cicadas within striking distance by mimicking the female part of the cicada mating duet - a trick requiring it to recognize complex patterns of sound and precisely when to chip in.Īwesome? Absolutely. Katydids - there are thousands of species - have the smallest ears of any animal, one on each front leg just below the “knee.” But their small size and seemingly strange location belie the sophisticated structure and impressive capabilities of these organs: to detect the ultrasonic clicks of hunting bats, pick out the signature songs of prospective mates, and home in on dinner. The story is told via comic-like segments that make it clear that the overall theme is humor-based.Kingdom Rush is a medieval tower defense series where your goal is to defend your kingdom against invading forces. If you’re looking for more tower defense, Kingdom Rush Vengeance has you covered. The combination of level and enemy variety, heroes who get progressively stronger, a long list of towers, and upgrades to tie it all together works well and few in the genre stick out as much in my mind as this one. As is normal for these titles, I only played a level or two at a time before I was satisfied with it for a bit, but I kept coming back for more. If you’re a fan of tower defense games with progression, this one is likely to keep your interest for a while. Kingdom Rush Vengeance is the latest of the series and it doesn’t disappoint. The only choice harder than decided which set of towers to bring in is which singular hero you’re going to have to supplement your defenses at any given time. Whether it’s a burly orc or a terrifying necromancer, your bases are covered. Excitingly enough, there are over a dozen of these, all of which have wildly varying strengths and weaknesses. Their mobility allows you to maneuver them where they’re needed and a plethora of special abilities lets them really bring the hurt to the enemy. Though you are only able to bring one into the fray, they’re incredibly powerful and can easily outmatch a handful of towers at higher levels. Heroes are also a powerful asset that you have at your disposal. You can only have five available to you during any single scenario though. There’s no shortage of towers for your to choose from. One tower may fire rapidly at single targets while another hits the entire area surrounding it and in each scenario you’ll have to decide which are best suited for the job. Attack speed, damage, status effects, and whether they do magical or physical damage are all factors in how effective they are against your foes. There’s a crazy number of these, each working in a different enough way that they’re exciting. Towers are your primary tool for wrecking and/or inconveniencing the assaulting force. Though the enemy is many and your forces are few, you’ve got several significant advantages that can swing things in your favor if you use them wisely. Each map is unique and you’ll find yourself visiting plenty of exciting places. The variety of enemies is impressive as well, with multiple factions to be encountered, each with at least half a dozen different units to complicate matters, including those that can fly, attack from a range, or freeze your defenders, to name a few. One map may be a desolate wasteland while another is a snow and ice covered tundra with ice that must be dealt with to build on the spots that it’s covering. Although those core components are always there, it’s amazing how much of a difference rearranging them or throwing in an additional element here or there can make. Totally not a boss fight.Įvery level is a surprisingly different experience in this title. Just ignore that dwarf seated on the bronze throne surrounded by gold. Fortunately, each dead enemy is a batch of coins in your pocket that can be used to further bolster your defenses. It’s easier said than done, especially when every once in a while you’ll deal with a boss that must be killed or you’ll lose and have to start the entire level over. You’re only allowed a certain number of enemies that make it from the entrance of their path to the exit and your goal is to kill them off before they can make their journey. Each level consists of paths that the enemies travel on, predetermined spots that you can build upon, and a variety of enemy types. As with most tower defense titles, Kingdom Rush Vengeance is made up of a series of levels. |
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